无法发送邮件 - javax.net.ssl.SSL异常：无法识别的SSL消息，纯文本连接？

我们正在使用Spring JavaMailSenderImpl发送邮件。以下是配置

 <bean id="mailSender" class="org.springframework.mail.javamail.JavaMailSenderImpl">
        <property name="host" value="${host}"/>
        <property name="port" value="${port}"/>
        <property name="username" value="${mail.username}"/>
        <property name="password" value="${mail.password}"/>
        <property name="javaMailProperties">
            <props>
                <!-- Use SMTP transport protocol -->
                <prop key="mail.transport.protocol" >${mail.transport.protocol}</prop>
                <!-- Use SMTP-AUTH to authenticate to SMTP server -->
                <prop key="mail.smtp.auth">${mail.smtp.auth}</prop>
                <!-- Use TLS to encrypt communication with SMTP server -->
                <prop key="mail.smtp.starttls.enable">${mail.smtp.starttls.enable}</prop>
                <prop key="mail.debug">false</prop>
                <prop key="mail.smtp.ssl.enable">true</prop>
            </props>
        </property>
    </bean>

属性文件 ：-

host=XXXX.XXXX.XX
port=25
mail.username=xxxxxxxx
mail.password=xxxxxxx
mail.transport.protocol=smtp
mail.smtp.auth=true
mail.smtp.starttls.enable=true

控制台日志

Exception in thread "taskExecutor-2" org.springframework.mail.MailSendException: Mail server connection failed; nested exception is javax.mail.MessagingException: Could not connect to SMTP host: XXXX.XXXX.XX, port: 25;
      nested exception is:
            javax.net.ssl.SSLException: Unrecognized SSL message, plaintext connection?. Failed messages: javax.mail.MessagingException: Could not connect to SMTP host: XXXX.XXXX.XX, port: 25;
      nested exception is:
            javax.net.ssl.SSLException: Unrecognized SSL message, plaintext connection?; message exception details (1) are:
    Failed message 1:
    javax.mail.MessagingException: Could not connect to SMTP host: XXXX.XXXX.XX, port: 25;
      nested exception is:
            javax.net.ssl.SSLException: Unrecognized SSL message, plaintext connection?
            at com.sun.mail.smtp.SMTPTransport.openServer(SMTPTransport.java:1934)
            at com.sun.mail.smtp.SMTPTransport.protocolConnect(SMTPTransport.java:638)
            at javax.mail.Service.connect(Service.java:295)
            at org.springframework.mail.javamail.JavaMailSenderImpl.doSend(JavaMailSenderImpl.java:389)
            at org.springframework.mail.javamail.JavaMailSenderImpl.send(JavaMailSenderImpl.java:340)
            at org.springframework.mail.javamail.JavaMailSenderImpl.send(JavaMailSenderImpl.java:336)
            at com.XXXX.Mailer$1.run(Mailer.java:52)
            at java.util.concurrent.ThreadPoolExecutor.runWorker(ThreadPoolExecutor.java:1145)
            at java.util.concurrent.ThreadPoolExecutor$Worker.run(ThreadPoolExecutor.java:615)
            at java.lang.Thread.run(Thread.java:744)
    Caused by: javax.net.ssl.SSLException: Unrecognized SSL message, plaintext connection?
            at sun.security.ssl.InputRecord.handleUnknownRecord(InputRecord.java:671)
            at sun.security.ssl.InputRecord.read(InputRecord.java:504)
            at sun.security.ssl.SSLSocketImpl.readRecord(SSLSocketImpl.java:927)
            at sun.security.ssl.SSLSocketImpl.performInitialHandshake(SSLSocketImpl.java:1312)
            at sun.security.ssl.SSLSocketImpl.startHandshake(SSLSocketImpl.java:1339)
            at sun.security.ssl.SSLSocketImpl.startHandshake(SSLSocketImpl.java:1323)
            at com.sun.mail.util.SocketFetcher.configureSSLSocket(SocketFetcher.java:507)
            at com.sun.mail.util.SocketFetcher.getSocket(SocketFetcher.java:238)
            at com.sun.mail.smtp.SMTPTransport.openServer(SMTPTransport.java:1900)
            ... 9 more

我们相信这与SSL证书无关，因为在同一台服务器中部署了其他Web应用程序，这些应用程序以相同的配置完美地发送电子邮件。这里可能有什么问题？